1. Abbasi, H., Naderi, R., Kafi, M., Azadi, P., Padasht Dahkaei, M.N. (2019). Interspecific hybridization between Lilium ledebourii and commercial cultivars of Lilium by cut-style method and ovary slice culture. Iranian Journal of Horticultural Science, 50(2), 287-294. (In Persian)

2. Alishah, O., Omidi, M. (2008). Laboratory Methods of Cytogenetic. Tehran University Press. 188 p. (In Persian).

3. Azadi, P., Khosh-Khui, M. (2007). Micropropagation of Lilium ledebourii Bioss. as affected by plant growth regulator, sucrose concentration, harvesting season and cold treatment. Electronic Journal of Biotechnology, 10, 583-591. [DOI:10.2225/vol10-issue4-fulltext-7]

4. Bakhshaie, M., Babalar, M., Mirmasoumi, M., Khalighi, A. (2010). Effects of light, sucrose, and cytokinins on somatic embryogenesis in Lilium ledebourii (Baker) Bioss. via transverse thin cell-layer cultures of bulblet microscales. The Journal of Horticultural Science and Biotechnology, 85(6), 491-496. [DOI:10.1080/14620316.2010.11512703]

5. Chi, H. S. (2002). The efficiencies of various embryo rescue methods in interspecific crosses of Lilium. Botanical Bulletin of Academia Sinica, 43, 139-146.

6. Chung, M. Y, Chung, J. D., Ramanna, M., van Tuyl J. M., Lim, K. B. (2013) Production of polyploids and unreduced gametes in Lilium auratum × L. henryi hybrid. Journal of Biological Sciences, 9(7), 693-701. [DOI:10.7150/ijbs.6427]

7. Du, X. Zhu, X., Yang, Y., Zhao, F., Liu, H. (2018). Pollen ultra-morphology and pollen viability test of Lilium Oriental hybrids. International Journal of Agriculture and Biology, 20(8), 1903-1907. [DOI:10.17957/IJAB/15.0753]

8. Erilova, A., Brownfield, L., Exner, V., Rosa, M., Twell, D., Mittelsten Scheid, O., Hennig, L., Köhler, C. (2009). Imprinting of the polycomb group gene MEDEA serves as a ploidy sensor in Arabidopsis. PLoS Genetics, 5(9):e1000663. [DOI:10.1371/journal.pgen.1000663]

9. Farsam, H., Amanlou, M., Amin, G., Nezamivand-Chegini, G., Salehi-Surmaghi, M. H., Shafiee, A. (2003). Anatomical and phytochemical study of Lilium ledebourii (Baker) Boiss., a rare endemic species in Iran. DARU Journal of Pharmaceutical Sciences, 11(4), 164-170.

10. He, G., Hu, F., Ming, J., Liu, C., Yuan, S. (2017). Pollen viability and stigma receptivity in Lilium during anthesis. Euphytica 213, 1-10. [DOI:10.1007/s10681-017-2019-9]

11. Ikeda, N., Niimi, Y., Han, D. S. (2003). Production of seedlings from ovules excised at the zygote stage in Lilium spp. Plant Cell, Tissue and Organ Culture, 73(2), 159-166. [DOI:10.1023/A:1022818413617]

12. Kim, J. Y., Song, Y. S., Na, J. K., Kim, J. H. (2022). Interspecific crossing between Lilium hansonii Leichtlin and L. brownii var. Colchesteri for the breeding of new lily cultivars. Agronomy, 12(3), 621. [DOI:10.3390/agronomy12030621]

13. Kwon, M. J., Ramzan, F., Ahn, Y. J., Hwang, Y. J., Kang, Y. I., Kim, C. K., Younis, A., Lim, K. B. (2017). Chromosomal analysis of Lilium longiflorum x Asiatic hybrids using GISH (genomic in situ hybridization). Horticulture, Environment, and Biotechnology, 58, 591-600. [DOI:10.1007/s13580-017-0019-2]

14. Marasek-Ciolakowska, A., Nishikawa, T., Shea, D. J., Okazaki, K. (2018). Breeding of lilies and tulips-Interspecific hybridization and genetic background. Breeding Science, 68(1), 35-52. [DOI:10.1270/jsbbs.17097]

15. Moradi Ashur B., Mohammadi R. Azimi M.H. (2010). Chromosome counting and ploidy level determination in different cultivars of Lilium. Agricultural Research, Education and Extension Organization (AREEO). No. 1059805 (In Persian).

16. Murashige, T. Skoog, F. (1962). A revised medium for rapid growth and bioassays with tobacco tissue cultures. Physiologia Plantarum, 15(3), 473-497. [DOI:10.1111/j.1399-3054.1962.tb08052.x]

17. Rechinger, K.H. (1989) Flora Iranica, no. 165, Liliaceae. Akademische Druck-u, Verlgsantalt, Graz.

18. Robinson, K.E.P. Firozabady, E. (1993) Transformation of floriculture crops. Scientia Horticulturae 55, 83-99. [DOI:10.1016/0304-4238(93)90026-M]

19. Saeedifard, M., Hosseini, S.M., Padasht Dehkaie, M.N. (2008). Modeling of the spatial distribution of the rare plant Lilium ledebourii. Rostaniha, 9(2), 137-150. (In Persian).

20. Sheikh-Assadi, M., Naderi, R., Kafi, M., Fatahi, R., Salami, S. A., Shariati, V. (2022). Complete chloroplast genome of Lilium ledebourii (Baker) Boiss and its comparative analysis: lights into selective pressure and adaptive evolution. Scientific Reports, 12(1), 1-17. [DOI:10.1038/s41598-022-13449-x]

21. Shokrollahi, S., Yousefzadeh, H., Parisod, C., Heshmati, G., Bina, H., Ali, S., Amirchakhmaghi, N., Song, Y. (2022). Phylogenetics and biogeography of Lilium ledebourii from the Hyrcanian forest. Diversity, 14, 137. [DOI:10.3390/d14020137]

22. Singh, R. J. (2016). Plant cytogenetics. CRC press. [DOI:10.1201/9781315374611]

23. Subasinghe Arachchige, E. C., Evans, L. J., Samnegård, U., Rader, R. (2022). Morphological characteristics of pollen from triploid watermelon and its fate on stigmas in a hybrid crop production system. Scientific Reports, 12(1), 1-12. [DOI:10.1038/s41598-022-06297-2]

24. Van Tuyl, J. M. De Jeu, M. J. (1997) Methods for Overcoming Interspecific Crossing Barriers. In: Shivanna, K-R., Sawhney, V-K. (eds), Pollen Biotechnology for Crop Production and Improvement. Cambridge University Press, pp 273-293. [DOI:10.1017/CBO9780511525469.015]

25. Van Tuyl, J. M., Arens, P., Shahin, A., Marasek-Ciolakowska, A., Barba-Gonzalez, R., Kim, H. T., et al. (2018). Lilium. In: J. van Huylenbroeck (ed), Ornamental Crops (Berlin: Springer), pp 481-512. [DOI:10.1007/978-3-319-90698-0_20]

26. Van Tuyl, J. M., Van Diën, M. P., Van Creij, M. G. M., Van Kleinwee, T. C. M., Franken, J. Bino, R. J. (1991). Application of in vitro pollination, ovary culture, ovule culture and embryo rescue for overcoming incongruity barriers in interspecific Lilium crosses. Plant Science, 74, 115-126. [DOI:10.1016/0168-9452(91)90262-7]

27. Wang, J., Huang, L., Bao, M. Z., Liu, G. F. (2009). Production of interspecific hybrids between Lilium longiflorum and L. lophophorum var. linearifolium via ovule culture at early stage. Euphytica 167, 45-55. [DOI:10.1007/s10681-008-9859-2]

28. Wendelbo, P. (1977) Tulips and Irises of Iran and Their Relatives. Botanical Institute of Iran, Tehran.

29. Xiao, K., Zheng, W., Zeng, J., Wu, L., Cui, L., Liu, Y., Yang, Y., Zhou, S. (2019). Analysis of abnormal meiosis and progenies of an odd-allotetraploid Lilium 'Honesty'. Scientia Horticulturae, 253, 316-321. [DOI:10.1016/j.scienta.2019.04.012]

30. Xiao, K., Zhu, Z., Zou, N., Zhang, L., Sun, Y., Zhou, S. (2022). The characteristics of abnormal meiosis and functional aneuploid pollen of odd-allotetraploid lily 'Honesty' unveiled using in situ hybridization. Scientia Horticulturae, 300, 111091. [DOI:10.1016/j.scienta.2022.111091]

31. Zhou, S., Tan, X., Fang, L., Jian, J., Xu, P., Yuan, G. (2013). Study of the female fertility of an odd-tetraploid of Lilium and its potential breeding significance. Journal of the American Society for Horticultural Science, 138(2), 114-119. [DOI:10.21273/JASHS.138.2.114]

32. Zhou, S., Zhou, G., Li, K. (2011). Euploid endosperm of triploid × diploid/tetraploid crosses results in aneuploid embryo survival in Lilium. HortScience, 46(4), 558-562. [DOI:10.21273/HORTSCI.46.4.558]

33. Abbasi, H., Naderi, R., Kafi, M., Azadi, P., Padasht Dahkaei, M.N. (2019). Interspecific hybridization between Lilium ledebourii and commercial cultivars of Lilium by cut-style method and ovary slice culture. Iranian Journal of Horticultural Science, 50(2), 287-294. (In Persian)

34. Alishah, O., Omidi, M. (2008). Laboratory Methods of Cytogenetic. Tehran University Press. 188 p. (In Persian).

35. Azadi, P., Khosh-Khui, M. (2007). Micropropagation of Lilium ledebourii Bioss. as affected by plant growth regulator, sucrose concentration, harvesting season and cold treatment. Electronic Journal of Biotechnology, 10, 583-591. [DOI:10.2225/vol10-issue4-fulltext-7]

36. Bakhshaie, M., Babalar, M., Mirmasoumi, M., Khalighi, A. (2010). Effects of light, sucrose, and cytokinins on somatic embryogenesis in Lilium ledebourii (Baker) Bioss. via transverse thin cell-layer cultures of bulblet microscales. The Journal of Horticultural Science and Biotechnology, 85(6), 491-496. [DOI:10.1080/14620316.2010.11512703]

37. Chi, H. S. (2002). The efficiencies of various embryo rescue methods in interspecific crosses of Lilium. Botanical Bulletin of Academia Sinica, 43, 139-146.

38. Chung, M. Y, Chung, J. D., Ramanna, M., van Tuyl J. M., Lim, K. B. (2013) Production of polyploids and unreduced gametes in Lilium auratum × L. henryi hybrid. Journal of Biological Sciences, 9(7), 693-701. [DOI:10.7150/ijbs.6427]

39. Du, X. Zhu, X., Yang, Y., Zhao, F., Liu, H. (2018). Pollen ultra-morphology and pollen viability test of Lilium Oriental hybrids. International Journal of Agriculture and Biology, 20(8), 1903-1907. [DOI:10.17957/IJAB/15.0753]

40. Erilova, A., Brownfield, L., Exner, V., Rosa, M., Twell, D., Mittelsten Scheid, O., Hennig, L., Köhler, C. (2009). Imprinting of the polycomb group gene MEDEA serves as a ploidy sensor in Arabidopsis. PLoS Genetics, 5(9):e1000663. [DOI:10.1371/journal.pgen.1000663]

41. Farsam, H., Amanlou, M., Amin, G., Nezamivand-Chegini, G., Salehi-Surmaghi, M. H., Shafiee, A. (2003). Anatomical and phytochemical study of Lilium ledebourii (Baker) Boiss., a rare endemic species in Iran. DARU Journal of Pharmaceutical Sciences, 11(4), 164-170.

42. He, G., Hu, F., Ming, J., Liu, C., Yuan, S. (2017). Pollen viability and stigma receptivity in Lilium during anthesis. Euphytica 213, 1-10. [DOI:10.1007/s10681-017-2019-9]

43. Ikeda, N., Niimi, Y., Han, D. S. (2003). Production of seedlings from ovules excised at the zygote stage in Lilium spp. Plant Cell, Tissue and Organ Culture, 73(2), 159-166. [DOI:10.1023/A:1022818413617]

44. Kim, J. Y., Song, Y. S., Na, J. K., Kim, J. H. (2022). Interspecific crossing between Lilium hansonii Leichtlin and L. brownii var. Colchesteri for the breeding of new lily cultivars. Agronomy, 12(3), 621. [DOI:10.3390/agronomy12030621]

45. Kwon, M. J., Ramzan, F., Ahn, Y. J., Hwang, Y. J., Kang, Y. I., Kim, C. K., Younis, A., Lim, K. B. (2017). Chromosomal analysis of Lilium longiflorum x Asiatic hybrids using GISH (genomic in situ hybridization). Horticulture, Environment, and Biotechnology, 58, 591-600. [DOI:10.1007/s13580-017-0019-2]

46. Marasek-Ciolakowska, A., Nishikawa, T., Shea, D. J., Okazaki, K. (2018). Breeding of lilies and tulips-Interspecific hybridization and genetic background. Breeding Science, 68(1), 35-52. [DOI:10.1270/jsbbs.17097]

47. Moradi Ashur B., Mohammadi R. Azimi M.H. (2010). Chromosome counting and ploidy level determination in different cultivars of Lilium. Agricultural Research, Education and Extension Organization (AREEO). No. 1059805 (In Persian).

48. Murashige, T. Skoog, F. (1962). A revised medium for rapid growth and bioassays with tobacco tissue cultures. Physiologia Plantarum, 15(3), 473-497. [DOI:10.1111/j.1399-3054.1962.tb08052.x]

49. Rechinger, K.H. (1989) Flora Iranica, no. 165, Liliaceae. Akademische Druck-u, Verlgsantalt, Graz.

50. Robinson, K.E.P. Firozabady, E. (1993) Transformation of floriculture crops. Scientia Horticulturae 55, 83-99. [DOI:10.1016/0304-4238(93)90026-M]

51. Saeedifard, M., Hosseini, S.M., Padasht Dehkaie, M.N. (2008). Modeling of the spatial distribution of the rare plant Lilium ledebourii. Rostaniha, 9(2), 137-150. (In Persian).

52. Sheikh-Assadi, M., Naderi, R., Kafi, M., Fatahi, R., Salami, S. A., Shariati, V. (2022). Complete chloroplast genome of Lilium ledebourii (Baker) Boiss and its comparative analysis: lights into selective pressure and adaptive evolution. Scientific Reports, 12(1), 1-17. [DOI:10.1038/s41598-022-13449-x]

53. Shokrollahi, S., Yousefzadeh, H., Parisod, C., Heshmati, G., Bina, H., Ali, S., Amirchakhmaghi, N., Song, Y. (2022). Phylogenetics and biogeography of Lilium ledebourii from the Hyrcanian forest. Diversity, 14, 137. [DOI:10.3390/d14020137]

54. Singh, R. J. (2016). Plant cytogenetics. CRC press. [DOI:10.1201/9781315374611]

55. Subasinghe Arachchige, E. C., Evans, L. J., Samnegård, U., Rader, R. (2022). Morphological characteristics of pollen from triploid watermelon and its fate on stigmas in a hybrid crop production system. Scientific Reports, 12(1), 1-12. [DOI:10.1038/s41598-022-06297-2]

56. Van Tuyl, J. M. De Jeu, M. J. (1997) Methods for Overcoming Interspecific Crossing Barriers. In: Shivanna, K-R., Sawhney, V-K. (eds), Pollen Biotechnology for Crop Production and Improvement. Cambridge University Press, pp 273-293. [DOI:10.1017/CBO9780511525469.015]

57. Van Tuyl, J. M., Arens, P., Shahin, A., Marasek-Ciolakowska, A., Barba-Gonzalez, R., Kim, H. T., et al. (2018). Lilium. In: J. van Huylenbroeck (ed), Ornamental Crops (Berlin: Springer), pp 481-512. [DOI:10.1007/978-3-319-90698-0_20]

58. Van Tuyl, J. M., Van Diën, M. P., Van Creij, M. G. M., Van Kleinwee, T. C. M., Franken, J. Bino, R. J. (1991). Application of in vitro pollination, ovary culture, ovule culture and embryo rescue for overcoming incongruity barriers in interspecific Lilium crosses. Plant Science, 74, 115-126. [DOI:10.1016/0168-9452(91)90262-7]

59. Wang, J., Huang, L., Bao, M. Z., Liu, G. F. (2009). Production of interspecific hybrids between Lilium longiflorum and L. lophophorum var. linearifolium via ovule culture at early stage. Euphytica 167, 45-55. [DOI:10.1007/s10681-008-9859-2]

60. Wendelbo, P. (1977) Tulips and Irises of Iran and Their Relatives. Botanical Institute of Iran, Tehran.

61. Xiao, K., Zheng, W., Zeng, J., Wu, L., Cui, L., Liu, Y., Yang, Y., Zhou, S. (2019). Analysis of abnormal meiosis and progenies of an odd-allotetraploid Lilium 'Honesty'. Scientia Horticulturae, 253, 316-321. [DOI:10.1016/j.scienta.2019.04.012]

62. Xiao, K., Zhu, Z., Zou, N., Zhang, L., Sun, Y., Zhou, S. (2022). The characteristics of abnormal meiosis and functional aneuploid pollen of odd-allotetraploid lily 'Honesty' unveiled using in situ hybridization. Scientia Horticulturae, 300, 111091. [DOI:10.1016/j.scienta.2022.111091]

63. Zhou, S., Tan, X., Fang, L., Jian, J., Xu, P., Yuan, G. (2013). Study of the female fertility of an odd-tetraploid of Lilium and its potential breeding significance. Journal of the American Society for Horticultural Science, 138(2), 114-119. [DOI:10.21273/JASHS.138.2.114]

64. Zhou, S., Zhou, G., Li, K. (2011). Euploid endosperm of triploid × diploid/tetraploid crosses results in aneuploid embryo survival in Lilium. HortScience, 46(4), 558-562. [DOI:10.21273/HORTSCI.46.4.558]

65. Abbasi, H., Naderi, R., Kafi, M., Azadi, P., Padasht Dahkaei, M.N. (2019). Interspecific hybridization between Lilium ledebourii and commercial cultivars of Lilium by cut-style method and ovary slice culture. Iranian Journal of Horticultural Science, 50(2), 287-294. (In Persian)

66. Alishah, O., Omidi, M. (2008). Laboratory Methods of Cytogenetic. Tehran University Press. 188 p. (In Persian).

67. Azadi, P., Khosh-Khui, M. (2007). Micropropagation of Lilium ledebourii Bioss. as affected by plant growth regulator, sucrose concentration, harvesting season and cold treatment. Electronic Journal of Biotechnology, 10, 583-591. [DOI:10.2225/vol10-issue4-fulltext-7]

68. Bakhshaie, M., Babalar, M., Mirmasoumi, M., Khalighi, A. (2010). Effects of light, sucrose, and cytokinins on somatic embryogenesis in Lilium ledebourii (Baker) Bioss. via transverse thin cell-layer cultures of bulblet microscales. The Journal of Horticultural Science and Biotechnology, 85(6), 491-496. [DOI:10.1080/14620316.2010.11512703]

69. Chi, H. S. (2002). The efficiencies of various embryo rescue methods in interspecific crosses of Lilium. Botanical Bulletin of Academia Sinica, 43, 139-146.

70. Chung, M. Y, Chung, J. D., Ramanna, M., van Tuyl J. M., Lim, K. B. (2013) Production of polyploids and unreduced gametes in Lilium auratum × L. henryi hybrid. Journal of Biological Sciences, 9(7), 693-701. [DOI:10.7150/ijbs.6427]

71. Du, X. Zhu, X., Yang, Y., Zhao, F., Liu, H. (2018). Pollen ultra-morphology and pollen viability test of Lilium Oriental hybrids. International Journal of Agriculture and Biology, 20(8), 1903-1907. [DOI:10.17957/IJAB/15.0753]

72. Erilova, A., Brownfield, L., Exner, V., Rosa, M., Twell, D., Mittelsten Scheid, O., Hennig, L., Köhler, C. (2009). Imprinting of the polycomb group gene MEDEA serves as a ploidy sensor in Arabidopsis. PLoS Genetics, 5(9):e1000663. [DOI:10.1371/journal.pgen.1000663]

73. Farsam, H., Amanlou, M., Amin, G., Nezamivand-Chegini, G., Salehi-Surmaghi, M. H., Shafiee, A. (2003). Anatomical and phytochemical study of Lilium ledebourii (Baker) Boiss., a rare endemic species in Iran. DARU Journal of Pharmaceutical Sciences, 11(4), 164-170.

74. He, G., Hu, F., Ming, J., Liu, C., Yuan, S. (2017). Pollen viability and stigma receptivity in Lilium during anthesis. Euphytica 213, 1-10. [DOI:10.1007/s10681-017-2019-9]

75. Ikeda, N., Niimi, Y., Han, D. S. (2003). Production of seedlings from ovules excised at the zygote stage in Lilium spp. Plant Cell, Tissue and Organ Culture, 73(2), 159-166. [DOI:10.1023/A:1022818413617]

76. Kim, J. Y., Song, Y. S., Na, J. K., Kim, J. H. (2022). Interspecific crossing between Lilium hansonii Leichtlin and L. brownii var. Colchesteri for the breeding of new lily cultivars. Agronomy, 12(3), 621. [DOI:10.3390/agronomy12030621]

77. Kwon, M. J., Ramzan, F., Ahn, Y. J., Hwang, Y. J., Kang, Y. I., Kim, C. K., Younis, A., Lim, K. B. (2017). Chromosomal analysis of Lilium longiflorum x Asiatic hybrids using GISH (genomic in situ hybridization). Horticulture, Environment, and Biotechnology, 58, 591-600. [DOI:10.1007/s13580-017-0019-2]

78. Marasek-Ciolakowska, A., Nishikawa, T., Shea, D. J., Okazaki, K. (2018). Breeding of lilies and tulips-Interspecific hybridization and genetic background. Breeding Science, 68(1), 35-52. [DOI:10.1270/jsbbs.17097]

79. Moradi Ashur B., Mohammadi R. Azimi M.H. (2010). Chromosome counting and ploidy level determination in different cultivars of Lilium. Agricultural Research, Education and Extension Organization (AREEO). No. 1059805 (In Persian).

80. Murashige, T. Skoog, F. (1962). A revised medium for rapid growth and bioassays with tobacco tissue cultures. Physiologia Plantarum, 15(3), 473-497. [DOI:10.1111/j.1399-3054.1962.tb08052.x]

81. Rechinger, K.H. (1989) Flora Iranica, no. 165, Liliaceae. Akademische Druck-u, Verlgsantalt, Graz.

82. Robinson, K.E.P. Firozabady, E. (1993) Transformation of floriculture crops. Scientia Horticulturae 55, 83-99. [DOI:10.1016/0304-4238(93)90026-M]

83. Saeedifard, M., Hosseini, S.M., Padasht Dehkaie, M.N. (2008). Modeling of the spatial distribution of the rare plant Lilium ledebourii. Rostaniha, 9(2), 137-150. (In Persian).

84. Sheikh-Assadi, M., Naderi, R., Kafi, M., Fatahi, R., Salami, S. A., Shariati, V. (2022). Complete chloroplast genome of Lilium ledebourii (Baker) Boiss and its comparative analysis: lights into selective pressure and adaptive evolution. Scientific Reports, 12(1), 1-17. [DOI:10.1038/s41598-022-13449-x]

85. Shokrollahi, S., Yousefzadeh, H., Parisod, C., Heshmati, G., Bina, H., Ali, S., Amirchakhmaghi, N., Song, Y. (2022). Phylogenetics and biogeography of Lilium ledebourii from the Hyrcanian forest. Diversity, 14, 137. [DOI:10.3390/d14020137]

86. Singh, R. J. (2016). Plant cytogenetics. CRC press. [DOI:10.1201/9781315374611]

87. Subasinghe Arachchige, E. C., Evans, L. J., Samnegård, U., Rader, R. (2022). Morphological characteristics of pollen from triploid watermelon and its fate on stigmas in a hybrid crop production system. Scientific Reports, 12(1), 1-12. [DOI:10.1038/s41598-022-06297-2]

88. Van Tuyl, J. M. De Jeu, M. J. (1997) Methods for Overcoming Interspecific Crossing Barriers. In: Shivanna, K-R., Sawhney, V-K. (eds), Pollen Biotechnology for Crop Production and Improvement. Cambridge University Press, pp 273-293. [DOI:10.1017/CBO9780511525469.015]

89. Van Tuyl, J. M., Arens, P., Shahin, A., Marasek-Ciolakowska, A., Barba-Gonzalez, R., Kim, H. T., et al. (2018). Lilium. In: J. van Huylenbroeck (ed), Ornamental Crops (Berlin: Springer), pp 481-512. [DOI:10.1007/978-3-319-90698-0_20]

90. Van Tuyl, J. M., Van Diën, M. P., Van Creij, M. G. M., Van Kleinwee, T. C. M., Franken, J. Bino, R. J. (1991). Application of in vitro pollination, ovary culture, ovule culture and embryo rescue for overcoming incongruity barriers in interspecific Lilium crosses. Plant Science, 74, 115-126. [DOI:10.1016/0168-9452(91)90262-7]

91. Wang, J., Huang, L., Bao, M. Z., Liu, G. F. (2009). Production of interspecific hybrids between Lilium longiflorum and L. lophophorum var. linearifolium via ovule culture at early stage. Euphytica 167, 45-55. [DOI:10.1007/s10681-008-9859-2]

92. Wendelbo, P. (1977) Tulips and Irises of Iran and Their Relatives. Botanical Institute of Iran, Tehran.

93. Xiao, K., Zheng, W., Zeng, J., Wu, L., Cui, L., Liu, Y., Yang, Y., Zhou, S. (2019). Analysis of abnormal meiosis and progenies of an odd-allotetraploid Lilium 'Honesty'. Scientia Horticulturae, 253, 316-321. [DOI:10.1016/j.scienta.2019.04.012]

94. Xiao, K., Zhu, Z., Zou, N., Zhang, L., Sun, Y., Zhou, S. (2022). The characteristics of abnormal meiosis and functional aneuploid pollen of odd-allotetraploid lily 'Honesty' unveiled using in situ hybridization. Scientia Horticulturae, 300, 111091. [DOI:10.1016/j.scienta.2022.111091]

95. Zhou, S., Tan, X., Fang, L., Jian, J., Xu, P., Yuan, G. (2013). Study of the female fertility of an odd-tetraploid of Lilium and its potential breeding significance. Journal of the American Society for Horticultural Science, 138(2), 114-119. [DOI:10.21273/JASHS.138.2.114]

96. Zhou, S., Zhou, G., Li, K. (2011). Euploid endosperm of triploid × diploid/tetraploid crosses results in aneuploid embryo survival in Lilium. HortScience, 46(4), 558-562. [DOI:10.21273/HORTSCI.46.4.558]

97. Abbasi, H., Naderi, R., Kafi, M., Azadi, P., Padasht Dahkaei, M.N. (2019). Interspecific hybridization between Lilium ledebourii and commercial cultivars of Lilium by cut-style method and ovary slice culture. Iranian Journal of Horticultural Science, 50(2), 287-294. (In Persian)

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